diff --git a/docs/CHANGELOG.txt b/docs/CHANGELOG.txt
index 07c882b..8d8ea0b 100644
--- a/docs/CHANGELOG.txt
+++ b/docs/CHANGELOG.txt
@@ -47,6 +47,8 @@
   glyph. (#2149, #515)
 - Selectable: Fixed honoring style.SelectableTextAlign with unspecified size. (#2347, #2601)
 - Selectable: Allow using ImGuiSelectableFlags_SpanAllColumns in other columns than first. (#125)
+- TreeNode: Made clicking on arrow with _OpenOnArrow toggle the open state on the Mouse Down
+  event rather than the Mouse Down+Up sequence (this is rather standard behavior).
 - Scrolling: Fixed scrolling centering API leading to non-integer scrolling values and initial
   cursor position. This would often get fixed after the fix item submission, but using the
   ImGuiListClipper as the first thing after Begin() could largely break size calculations. (#3073)
diff --git a/imgui_widgets.cpp b/imgui_widgets.cpp
index 3a360b9..08b09f1 100644
--- a/imgui_widgets.cpp
+++ b/imgui_widgets.cpp
@@ -5319,30 +5319,39 @@
         return is_open;
     }
 
-    // Flags that affects opening behavior:
-    // - 0 (default) .................... single-click anywhere to open
-    // - OpenOnDoubleClick .............. double-click anywhere to open
-    // - OpenOnArrow .................... single-click on arrow to open
-    // - OpenOnDoubleClick|OpenOnArrow .. single-click on arrow or double-click anywhere to open
-    ImGuiButtonFlags button_flags = 0;
+    ImGuiButtonFlags button_flags = ImGuiTreeNodeFlags_None;
     if (flags & ImGuiTreeNodeFlags_AllowItemOverlap)
         button_flags |= ImGuiButtonFlags_AllowItemOverlap;
-    if (flags & ImGuiTreeNodeFlags_OpenOnDoubleClick)
-        button_flags |= ImGuiButtonFlags_PressedOnDoubleClick | ((flags & ImGuiTreeNodeFlags_OpenOnArrow) ? ImGuiButtonFlags_PressedOnClickRelease : 0);
-    else
-        button_flags |= ImGuiButtonFlags_PressedOnClickRelease;
     if (!is_leaf)
         button_flags |= ImGuiButtonFlags_PressedOnDragDropHold;
 
     // We allow clicking on the arrow section with keyboard modifiers held, in order to easily
     // allow browsing a tree while preserving selection with code implementing multi-selection patterns.
     // When clicking on the rest of the tree node we always disallow keyboard modifiers.
-    const float hit_padding_x = style.TouchExtraPadding.x;
-    const float arrow_hit_x1 = (text_pos.x - text_offset_x) - hit_padding_x;
-    const float arrow_hit_x2 = (text_pos.x - text_offset_x) + (g.FontSize + padding.x * 2.0f) + hit_padding_x;
-    if (window != g.HoveredWindow || !(g.IO.MousePos.x >= arrow_hit_x1 && g.IO.MousePos.x < arrow_hit_x2))
+    const float arrow_hit_x1 = (text_pos.x - text_offset_x) - style.TouchExtraPadding.x;
+    const float arrow_hit_x2 = (text_pos.x - text_offset_x) + (g.FontSize + padding.x * 2.0f) + style.TouchExtraPadding.x;
+    const bool is_mouse_x_over_arrow = (g.IO.MousePos.x >= arrow_hit_x1 && g.IO.MousePos.x < arrow_hit_x2);
+    if (window != g.HoveredWindow || !is_mouse_x_over_arrow)
         button_flags |= ImGuiButtonFlags_NoKeyModifiers;
 
+    // Open behaviors can be altered with the _OpenOnArrow and _OnOnDoubleClick flags.
+    // Some alteration have subtle effects (e.g. toggle on MouseUp vs MouseDown events) due to requirements for multi-selection and drag and drop support.
+    // - Single-click on label = Toggle on MouseUp (default)
+    // - Single-click on arrow = Toggle on MouseUp (when _OpenOnArrow=0)
+    // - Single-click on arrow = Toggle on MouseDown (when _OpenOnArrow=1)
+    // - Double-click on label = Toggle on MouseDoubleClick (when _OpenOnDoubleClick=1)
+    // - Double-click on arrow = Toggle on MouseDoubleClick (when _OpenOnDoubleClick=1 and _OpenOnArrow=0)
+    // This makes _OpenOnArrow have a subtle effect on _OpenOnDoubleClick: arrow click reacts on Down rather than Up.
+    // It is rather standard that arrow click react on Down rather than Up and we'd be tempted to make it the default 
+    // (by removing the _OpenOnArrow test below), however this would have a perhaps surprising effect on CollapsingHeader()? 
+    // So right now we are making this optional. May evolve later.
+    if (is_mouse_x_over_arrow && (flags & ImGuiTreeNodeFlags_OpenOnArrow))
+        button_flags |= ImGuiButtonFlags_PressedOnClick;
+    else if (flags & ImGuiTreeNodeFlags_OpenOnDoubleClick)
+        button_flags |= ImGuiButtonFlags_PressedOnDoubleClick;
+    else
+        button_flags |= ImGuiButtonFlags_PressedOnClickRelease;
+
     bool selected = (flags & ImGuiTreeNodeFlags_Selected) != 0;
     const bool was_selected = selected;
 
@@ -5356,7 +5365,7 @@
             if ((flags & (ImGuiTreeNodeFlags_OpenOnArrow | ImGuiTreeNodeFlags_OpenOnDoubleClick)) == 0 || (g.NavActivateId == id))
                 toggled = true;
             if (flags & ImGuiTreeNodeFlags_OpenOnArrow)
-                toggled |= (g.IO.MousePos.x >= arrow_hit_x1 && g.IO.MousePos.x < arrow_hit_x2) && (!g.NavDisableMouseHover); // Lightweight equivalent of IsMouseHoveringRect() since ButtonBehavior() already did the job
+                toggled |= is_mouse_x_over_arrow && !g.NavDisableMouseHover; // Lightweight equivalent of IsMouseHoveringRect() since ButtonBehavior() already did the job
             if ((flags & ImGuiTreeNodeFlags_OpenOnDoubleClick) && g.IO.MouseDoubleClicked[0])
                 toggled = true;
             if (g.DragDropActive && is_open) // When using Drag and Drop "hold to open" we keep the node highlighted after opening, but never close it again.